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Bio Med Central Open Access Page 1 of 10(page number not for citation purposes)Biology Direct Research Inferring clocks when lacking rocks: the variable rates of molecular evolution in bacteria Chih-Horng Kuo and Howard Ochman*Address: Department of Ecology & Evolutionary Biology, University of Arizona, Tucson, Arizona 85721 USAEmail: Chih-Horng Kuo - email@example.com; Howard Ochman* - firstname.lastname@example.org* Corresponding author Abstract Background: Because bacteria do not have a robust fossil record, attempts to infer the timing ofevents in their evolutionary history requires comparisons of molecular sequences.
This use ofmolecular clocks is based on the assumptions that substitution rates for homologous genes or sitesare fairly constant through time and across taxa.
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For these sites, we show estimated metrics based on traffic patterns across the web as a whole.Genetic drift plays a fundamental role in determining the accumulation ofsubstitutions in 16S r RNA genes and at nonsynonymous sites.Moreover, divergence estimatesbased on a set of universally conserved protein-coding genes also exhibit low correspondence tothose based on 16S r RNA genes.Certified Site Metrics are metrics that are directly-measured from the website instead of estimated.The website owner has installed an Alexa Certify Code on the pages of their site and chosen to show the metrics publicly.